Clypeoloculus Kaz. Tanaka & K. Hiray., in Tanaka et al., Stud. Mycol. 82: 108 (2015).
Index Fungorum number: IF 811318; Facesoffungi number: FoF 08307, 11 morphological species (Species Fungorum, 2022), 11 species with molecular data.
Saprobic on host. Sexual morph: Ascomata scattered to 2–3 grouped, immersed to erumpent, subglobose or hemispherical with flattened base in section, covered with numerous brown hyphae around ascomata. Ostiolar neck terete to papillate, central, clypeate, composed of carbonaceous cells. Ascomatal wall wedge-shaped or “rim-like” at sides. Pseudoparaphyses septate, branched and anastomosed. Asci fissitunicate, clavate, with a short stipe, 8-spored. Ascospores narrowly fusiform, slightly curved, with a primary septum median to supramedian, constricted at the septum, hyaline, smooth, surrounded by an entire gelatinous sheath. Asexual morph: unknown (Adapted from Tanaka et al., 2015).
Type species: Clypeoloculus akitaensis Kaz. Tanaka & K. Hiray.
Notes: Clypeoloculus was introduced by Tanaka et al. (2015) with C. akitaensis as type species. Clypeoloculus is characterized by immersed to erumpent, subglobose ascomata, terete to papillate ostiolar neck, clavate asci and narrowly fusiform, slightly curved ascospores surrounded by a gelatinous sheath. Clypeoloculus morphologically resembles Massarina s. str. but differs in having ascomata with wedge shaped or “rim-like” ascomatal wall and narrowly fusiform ascospores (Aptroot, 1998; Tanaka & Harada, 2003b). Clypeoloculus is phylogenetically distinct from Massarina as it belongs to Morosphaeriaceae while Massarina belongs to Massarinaceae. Clypeoloculus is morphologically and phylogenetically a distinct genus in Morosphaeriaceae. Molecular markers available for Clypeoloculus include LSU, SSU, ITS and TEF1.
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