Phacellium
Phacellium Bonord., in Rabenhorst, Fungi europ. exsicc. Edn 2, ser. 2: no. 288 (1860).
Index Fungorum number: IF 9286; Facesoffungi number: FoF 08637, 13 morphological species (Species Fungorum 2022), 1 species with molecular data.
Associated with leaf spots. Sexual morph: Unknown. Asexual morph: Leaf spots amphigenous, subcircular to angular-irregular, occasionally confluent, pale to dark brown, later grayish brown to dingy gray, on the lower leaf surface usually much paler, margin conspicuous, narrow, dark brown to blackish. Mycelium internal, forming immersed stromatic hyphal aggregations at the base of synnemata, brown. Conidiomata synnematous, erect, straight, occasionally slightly curved to sinous, subcylindrical or somewhat attenuated toward s the apex, medium brown, composed of a firm stipe of densely appressed parallel threads and a rather compact, often poorly developed capitulum, barely or only slightly splaying out. Individual conidiophores, pale to mid olivaceous-brown, pluriseptate throughout smooth, thin-walled. Conidiogenous cells integrated, terminal, intercalary and pleurogenous, subhyaline to very pale olivaceous or olivaceous brown, with a single to usually several conspicuous conidiogenous loci, slightly thickened and darkened. Conidia catenate, in simple or occasionally branched chains, ellipsoid-ovoid, cylindrical, 0–1(–2)-septate, hyaline or subhyaline, thin-walled, smooth or almost so, ends obtuse to somewhat attenuated, hila slightly thickened and darkened (adapted from description of Phacellium annonae-cherimoliae in Braun & Crous, 2008).
Type species: Phacellium inhonestum Bonord.
Notes: Braun (1990) discussed the taxonomy of Phacellium and Isariopsis and re-corrected the older, valid name Phacellium. Braun (1998) provided a monograph of Phacellium and accepted 22 species. Braun (1998) listed Mycosphaerella pseudomaculiformis as the sexual morph of P. bulbigerum. Crous et al. (2011) considered Phacellium as a synonym of Ramularia based on R. pusilla. The placement of Phacellium in Mycosphaerellaceae is doubtful. Only ITS sequence data is available for Phacellium veronicae. More taxa with DNA sequence data are needed to confirm the taxonomic position of Phacellium.
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