Dothideomycetes » Dothideomycetes, genera incertae sedis


Citation: Pem D et al. (2019) Mycosphere Notes 275-324: A morphotaxonomic revision and typification of obscure Dothideomycetes genera (incertae sedis). Mycosphere 10(1), 1115–1246


Achorella Theiss. & Syd. Annls mycol. 13(3/4): 340 (1915)

Parasitic or epiphytic on the surface of living leaves. Sexual morph: Undetermined. Asexual morph: Conidiomata pycnidial, eustromatic, superficial, multilocular, black, subglobose, aggregated, thick-walled; walls composed of dark brown textura angularis. Conidiophores hyaline, cylindrical, reduced to conidiogenous cells. Conidiogenous cells phialidic, cylindrical, 1-septate. Conidia hyaline, 2-celled, septate, cylindrical, apical obtuse, smooth-walled with tiny setula or appendage.


Type species Achorella ametableta (Rehm).


Notes – The genus Achorella was described and introduced by Theissen and Sydow (1915) with A. ametableta as the type species. Achorella ametableta was found on Baccharis (Asteraceae) stems in Brazil and is characterized by globose locules with the perithecial protuberances immersed in the stroma. Locules are spherical to pear-shaped with a short neck. Paraphyses are numerous, thin, cylindrical and filamentous. Ascospores are spindle shaped-constricted on both sides tapered in the middle, brown. It was earlier placed in the Dothideae, within the order Dothideales (Clement 1931), but is now in Dothideomycetes, genera incertae sedis (Wijayawardene et al. 2018). There are currently ten species in this genus. Cultures and sequences are unavailable. The growth of superficial mycelium along with the free perithecium-like protuberances are similar to characters of the species in the Dothideales and of the Perisporiaceae though its closest relation is with the former. The specimen that we observed was in poor condition hence we treat this taxa as doubtful until the fungus is recollected.



Clements FE, Shear CL. 1931 – The genera of Fungi. H.W. Wilson Co., New York pp 1–496.

Theissen F, Sydow H. 1915 – Die Dothideales. Kritisch-systematische Originaluntersuchungen. Annales Mycologici. 13(3-4), 147–746.

Wijayawardene NN, Hyde KD, Lumbsch HT, Liu JK et al. 2018 – Outline of Ascomycota: 2017. Fungal Diversity 88, 167–263.


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